Since the School’s first graduating class in 2004, 145 students have been awarded doctoral degrees from the School. The overall retention rate is 95%. Of those earning a degree: 89% earned a Ph.D.; 11% graduated with a Master’s degree.

All students complete their degree within 6 years. The average time to degree is 5.19 years.

Students have published almost 500 papers, a demonstration of the rigorous and reproducible work they performed during their training. About a quarter of the students have successfully applied for external fellowships.

A third of our graduates are in faculty positions, most in tenure-track positions at major U.S or foreign research institutions, and a third in industry. Other graduates are in academic research, postdoctoral research, or other scientific careers, such as university administrators, CEOs of biotech companies, and consultants.

Ph.D. Student Progress & Time to Degree

Career Paths of Ph.D. Students

Graduates in Faculty or Independent Positions (in order of completion)

Name Faculty Position
Ira Hall Professor, Yale University, CT
Niraj Tolia Senior Investigator, NIAID, National Institutes of Health
Patrick Paddison Professor, Fred Hutchinson Cancer Research Center, WA
Elizabeth Bartom (nee Thomas) Assistant Professor, Northwestern University, IL
Michelle Heck (nee Cilia) Research Molecular Biologist, USDA-ARS and Adjunct Associate Professor, Cornell University
Zachary Lippman Professor, Cold Spring Harbor Laboratory & Investigator, Howard Hughes Medical Institute
Ji-Joon Song Professor, Korea Advanced Institute of Science and Technology (KAIST), South Korea
Elena Ezhkova Associate Professor, Mount Sinai School of Medicine, NY
Masafumi Muratani Professor, University of Tsukuba, Japan
Santanu Chakraborty Program Director for Sustainable Life Sciences and Associate Professor, Atria University, India
Claudia Feierstein Research Associate, Champalimaud Neuroscience Programme, Portugal
Gowan Tervo Group Leader, HHMI, Janelia Research Campus, VA
Marco Mangone Professor, Arizona State University, AZ
Elizabeth Murchison Professor, Cambridge University, UK
Hiroki Asari Group leader, EMBL Monterotondo, Rome
François Bolduc Associate Professor, University of Alberta, Canada
Wei Wei Professor, University of Chicago, IL
Christopher Harvey Associate Professor, Harvard University, MA
Tomas Hromadka Group Leader, Institute of Neuroimmunology, Slovak Academy of Sciences
Monica Dus Associate Professor, University of Michigan, MI and White House Fellow, United States Department of the Navy
Shu-Ling Chiu Assistant Professor, Academia Sinica, Taiwan
Daniel Chitwood Assistant Professor, Michigan State University, MI
Jeremy Wilsuz Associate Professor, Baylor College of Medicine, TX
Shraddha Pai Investigator, Ontario Institute for Cancer Research, Toronto
Oliver Fregoso Assistant Professor, University of California, Los Angeles, CA
Hiroshi Makino Assistant Professor, Nanyang Technological University, Singapore
Galen Collins Assistant Professor, Mississippi State University, MS
Katherine McJunkin Stadtman Tenure Track Investigator, National Institutes of Health
Zhenxun Wang Adjunct Junior Principal Investigator, Singapore Eye Research Institute and Adjunct Assistant Professor Duke-NUS Medical School, Singapore
Eyal Gruntman Assistant Professor, University of Toronto Scarborough, Canada
Petr Znamenskiy Assistant Professor, The Crick Institute, UK
Yevgeniy Plavskin Visiting Clinical Associate Professor, New York University
Michael Pautler Head, Genomics Services, Platform Genetics & Vineland Research and Innovation Centre, Canada
Melanie Eckersley-Maslin Assistant Professor, Peter Mac Cancer Centre at University of Melbourne
Philippe Batut Assistant Professor, Columbia University (January 2024)
Elvin Wagenblast Assistant Professor, Mount Sinai School of Medicine, NY
Kristen Delevich Assistant Professor, Washington State University
Cinthya Zepeda Mendoza Medical Director, Cytogenetics and Genomic Microarray at ARUP Laboratories. Assistant Professor, Department of Pathology, University of Utah
Wee Siong Goh Assistant Professor Shenzhen Bay Laboratory, China
Arkarup Bandyopadhyay Assistant Professor, Cold Spring Harbor Laboratory, NY
Charles Underwood Professor, Radboud University, The Netherlands
Justus Kebschull Assistant Professor, The Johns Hopkins University
Paul Masset Assistant Professor, McGill University, Canada (January 2024)
Brianna Bibel Visiting Professor, Saint Mary’s College of California

Graduates in Industry Positions (in order of completion)

Student Current Position
Amy Caudy Principal, Scientific Operations, Maple Flavored Solutions, NY*
Emiliano Rial-Verde Vice President, Food & Ingredients Strategy, Bunge Limited, NY
Rebecca Ewald International Business Leader, Ventana Medical Systems/Roche, AZ
Catherine Seiler (nee Cormier) Director, Operational Excellence in Translation Medicine, AstraZeneca, MA
Allison Blum Senior Director, Scientific Affairs, Pfizer Research and Development, NY
Yaniv Erlich CEO, Eleven Therapeutics, UK*
Colin Malone Co-Founder & Head of Biology at VNV NewCo, NY*
Amy Leung Principal Solution Engineer, DNAnexus, CA
Amy Rappaport Senior Director, Translational Immunology, Gritstone bio, Inc., CA
Claudio Scuoppo Principal Scientist, Sapience Therapeutics, Inc.
Frederick Rollins Director, Competitive Intelligence, Oncology R&D, AstraZeneca, Washington DC
Patrick Finigan Associate Director, Regulatory Affairs CMC, Gilead Sciences
Kyle Honegger Data Scientist, Ann & Robert H. Lurie Children’s Hospital of Chicago, IL
Maria Pineda Co-Founder, CEO, Envisagenics, NY
Felix Schlesinger Bioinformatics Lead, Scale Bio, CA
Megan Hogan (nee Bodnar) Team Lead, Neochromosome, Inc., NY
Paloma Guzzardo Head, Target Biology, Variant Bio, WA
Saya Ebbesen Associate Director, Medical + Scientific Strategy at BluPrint Oncology, London, UK
Hassana Oyibo Associate Director, Synthetic Biology, Tessera Therapeutics
Michael Pautler Head, Genomics Services, Platform Genetics, Canada*
Marek Kudla Director of Bioinformatics, Ardigen, Poland
Joshua Sanders Founder and C.E.O., Sanworks, L.L.C., NY
Katie Liberatore Molecule Design Director, Cibus, MN
Kaja Wasik Co-Founder and Chief Scientific Officer, Gencove & Variant Bio, NY
Stephane Castel Co-Founder and Chief Technology Officer, Variant Bio, WA
Mitchell Bekritsky Senior Manager, Bioinformatics, Illumina, Inc.,
Sang-Geol Koh Scientist and Entrepreneur, {Mind}, South Korea
Ozlem Aksoy (nee Mert) Senior Scientist, Pfizer, CA
Susann Manchado (nee Weissmueller) Strategic Assistant (Chief of Staff) for President Global Drug Development, Novartis, Switzerland
Nilgun Tasdemir Senior Scientist, Pfizer, NY
Silvia Fenoglio Senior Scientist, Tango Therapeutics, MA
Jack Walleshauser Senior Scientist, Spotlight Therapeutics, CA
Lisa Krug Senior Scientist, Kallyope, NY
Robert Aboukhalil Senior Software Engineer, Chan Zuckerberg Initiative
Anja Hohmann Director, Cell Therapies at Be Biopharma, MA
Matt Koh Research Engineer, ASAAP, NY
Annabel Romero Hernandez Director of AI for Drug Discovery and Molecular Modeling, SFL Scientific, MA
Maria Nattestad Senior Software Engineer, Google, CA
Onyekachi Odoemene Research Scientist, ML Solutions Lab, Amazon
Daniel Kepple Engineer, Meta, NY
Sashank Pisupati Machine Learning Scientist, Limbic, UK
Kristina Grigaityte Associate Director, Data Science, Merck
Elizabeth Hutton Senior Computational Biologist, ArtisanBio, CO
Ngoc (Tumi) Tran Machine Learning Scientist, Relay Therapeutics, MA
Katie Meze Senior Scientist, Structural Biology, Sanofi, MA
Benjamin Berube Research Scientist, Monocot Breeding and Biotechnology, Ohalo Genetics, CA
Shaina Lu Senior Data Scientist, R&D, Miltenyi Biotec
Benjamin Harris Computational Biologist II, Lyell Immunopharma, CA
Christopher Krasniak Research Data Scientist, Somatus, NH
Ran (Rena) Yan Senior Scientist, Parthenon Therapeutics, MA
Martyna Sroka Scientist, Relay Therapeutics, MA
Sara Boyle Data Analytics Consultant, Northern Trust, IL
Asad Aziz Lakhani Associate, Search and Evaluation, Autobahn Labs

*Left a faculty position

Graduates in Postdoctoral or Academic Research Positions (in order of completion)

Student Current Position
Charles Kopec Associate Professional Specialist, Princeton University (Advisor, Dr. Carlos Brody)
Oliver Tam Research Scientist, NYU Institute for Systems Genetics, NY
Elizabeth Nakasone Junior Researcher, Translational and Clinical Research, University of Hawaiʻi Cancer Center
Claudio Scuoppo Instructor, Columbia University (Advisor: Riccardo Dalla-Favera)
Zhenxun Wang Research Fellow, Genome Institute of Singapore with (Advisor: Wai Leong Tam)
Eyal Gruntman Research Scientist, Janelia Farm (Advisor: Michael Reiser)
Yevgeniy Plavskin Postdoctoral Fellow, New York University (Advisor: Mark Siegal)
Zinaida Perova Project Lead, Cancer Models, European Bioinformatics Institute (EMBL-EBI), Cambridge, UK
Dario Bressan Head of IMAXT laboratory, CRUK, Cambridge Institute, UK
Philippe Batut Postdoctoral Fellow, Princeton University (Advisor: Michael Levine)
Joaquina Delas Vives Postdoctoral Fellow, Francis Crick Institute, UK (Advisor: James Briscoe)
Abram Santana Postdoctoral Fellow, Harvard University (Advisor: Joan Brugge)
Fred Marbach Senior Laboratory Research Scientist, Francis Crick Institute, UK (Advisor: Andreas Schaefer)
Yu-Jui (Ray) Ho Bioinformatic Specialist II, Memorial Sloan-Kettering Cancer Center (Advisor: Scott Lowe)
Paul Masset Postdoctoral Fellow, Harvard University (Advisor: Venkatesh Murthy and Naoshige Uchida)
Talitha Forcier Postdoctoral Fellow, New York University (Advisor: Molly Hammell)
Laura Maiorino Postdoctoral Fellow, Koch Institute, MIT (Advisor: Darrell Irvine)
Giorgia Battistoni Research Associate, Cancer Research UK (Advisor: Gregory Hannon)
Jacqueline Giovanniello Postdoctoral Fellow, UCLA (Advisor: Kate Wassum)
Hamza Giaffar Postdoctoral Fellow, UCSD (Advisor: Mikio Aoi)
Anqi Zhang Postdoctoral Fellow, Harvard University (Advisor: Florian Engert)
Alexandra Nowlan Postdoctoral Fellow, UNC Chapel Hill (Advisor: Zoe McElligott)
Katie Meze Postdoctoral Fellow, University of Michigan (Advisor: Nils Walters)
Christopher Krasniak Data Science Fellow, The Data Incubator
Luqun Shen Postdoctoral Fellow, Cornell University (Advisor: Alex Kwan)
Alberto Corona Postdoctoral Fellow, Icahn School of Med at Mount Sinai (Advisor: Paul Kenny)
Kathryn O’Neill Child Health Equity Postdoc, University of Washington, Seattle
David Johnson Postdoctoral Fellow, Invent at Seattle Children’s Postdoctoral Scholars Program (Advisor: Siobhan Pattwell)
Bruno Gegenhuber Postdoctoral Fellow, Harvard University (Advisor: Michael Greenberg)
Lyndsey Aguirre Biological Sciences Tech, Peanuts and Small Grains Research Unity, USDA-ARS, OK
Amritha Varshini Hanasoge Somasundara Senior Scientist, Memorial Sloan-Kettering Cancer Center (Advisor: Ross Levine)
Chengxiang (Charlie) Yuan Postdoctoral Fellow, Institute of Molecular and Cell Biology, Singapore (Advisor: Weimiao Yu)
Diogo Maia e Silva Postdoctoral Fellow, Mass General Hospital (Advisor: Keith Joung)
Yuzhao (Richard) Hu Postdoctoral Scientist, Corteva Agriscience, IA

Graduates in non-Research Positions (in order of completion)

Name Current Position
Ahmet M. Denli Associate Editor, Genome Research, CSHL, NY
Beth Chen Operations Manager at Homer Scientific Holdings Inc.
Darren Burgess Senior Editor, Nature Reviews Genetics, UK
Rebecca Bish-Cornelissen Head of Discovery and Preclinical Research, The Mark Foundation for Cancer Research, NY
Angelique Girard Director of Finance and Administration, Amplitude Studios, Paris, France
Allison Blum Managing Director, Scientific Communications, Life Sci Advisors, NY
Keisha John Associate Dean for Diversity, Equity and Inclusion, University of Virginia, VA
David Simpson Freelance Data Scientist and Strategy Consultant
Ian Peikon Co-founder & CSO, Cajal Neuroscience; Venture Partner, Lux Capital
Colleen Carlston Clinical Fellow in Pediatrics, Boston Children’s Hospital, MA
Tyler Garvin VP Operations, Underground Cellar, CA
Brittany Cazakoff Associate Lawyer, Cooley, CA
Lital Chartarifsky Associate Medical Director, FCB CURE, NY
Michael Gutbrod Senior Associate, Santé Ventures, MA
Emilis Bruzas Associate Director, Medical Strategy, FCB Health, NY
Sanchari Ghosh Content Specialist for BioRxiv, NY
Georgi Yordanov Equity Research Associate, Cowen and Company, NY
Jue Xiang Wang Consultant, Boston Consulting Group, NY
Matt Lee Oncology Medical Science Liaison, EMD Serono, Inc.
Sofya Polyanskaya Consultant, Scitaris, Berlin, Germany
Jenelys Ruiz Ortiz Bioinformatics Analyst, Bioinformatics Resource Center, The Rockefeller University, NY
Ilgin Ergin Foundation and Corporate Relations Coordinator, CSHL, NY
Alexa Pagliaro Foundation and Corporate Relations Officer, Whitehead Institute, MA