Since the School’s first graduating class in 2004, 127 students have been awarded doctoral degrees from the School. The overall retention rate is 95%. Of those earning a degree: 88% earned a Ph.D.; 12% graduated with a Master’s degree.

All students complete their degree within 6 years. The average time to degree is 5.17 years.

Students have published almost 500 papers, a demonstration of the rigorous and reproducible work they performed during their training. About a quarter of the students have successfully applied for external fellowships.

A third of our graduates are in faculty positions, most in tenure-track positions at major U.S or foreign research institutions, and a third in industry. Other graduates are in academic research, postdoctoral research, or other scientific careers, such as university administrators, CEOs of biotech companies, and consultants.

Ph.D. Student Progress & Time to Degree

Career Paths of Ph.D. Students

Graduates in Faculty or Independent Positions (in order of completion)

Name Faculty Position
Ira Hall Professor, Yale University, CT
Niraj Tolia Senior Investigator, NIAID, National Institutes of Health
Patrick Paddison Associate Member, Fred Hutchinson Cancer Research Center, WA
Elizabeth Bartom (nee Thomas) Assistant Professor, Northwestern University, IL
Michelle Heck (nee Cilia) Research Molecular Biologist, USDA-ARS and Adjunct Associate Professor, Cornell University
Zachary Lippman Professor, Cold Spring Harbor Laboratory & Investigator, Howard Hughes Medical Institute
Ji-Joon Song Professor, Korea Advanced Institute of Science and Technology (KAIST), South Korea
Elena Ezhkova Associate Professor, Mount Sinai School of Medicine, NY
Masafumi Muratani Professor, University of Tsukuba, Japan
Santanu Chakraborty Program Director for Sustainable Life Sciences and Associate Professor, Atria University, India
Claudia Feierstein Research Associate, Champalimaud Neuroscience Programme, Portugal
Gowan Tervo Laboratory Head & Principal Scientist, HHMI, Janelia Farm Research Campus, VA
Marco Mangone Associate Professor, Arizona State University, AZ
Elizabeth Murchison Professor, Cambridge University, UK
Hiroki Asari Group leader, EMBL Monterotondo, Rome
François Bolduc Associate Professor, University of Alberta, Canada
Wei Wei Associate Professor, University of Chicago, IL
Christopher Harvey Associate Professor, Harvard University, MA
Tomas Hromadka Group Leader, Institute of Neuroimmunology, Slovak Academy of Sciences
Monica Dus Assistant Professor, University of Michigan, MI
Shu-Ling Chiu Assistant Professor, Academia Sinica in Taiwan
Daniel Chitwood Assistant Professor, Michigan State University, MI
Jeremy Wilsuz Associate Professor, Baylor College of Medicine, TX
Shraddha Pai Investigator, Ontario Institute for Cancer Research, Toronto
Oliver Fregoso Assistant Professor, University of California, Los Angeles, CA
Amy Leung Assistant Research Professor, City of Hope’s Beckman Research Institute
Hiroshi Makino Assistant Professor, Nanyang Technological University, Singapore
Katherine McJunkin Stadtman Tenure Track Investigator, National Institutes of Health
Zhenxun Wang Adjunct Junior Principal Investigator, Singapore Eye Research Institute and Adjunct Assistant Professor Duke-NUS Medical School, Singapore
Petr Znamenskiy Assistant Professor, The Crick Institute, UK
Michael Pautler Head, Genomics Services, Platform Genetics & Vineland Research and Innovation Centre, Canada
Melanie Eckersley-Maslin Assistant Professor, Peter Mac Cancer Centre at University of Melbourne
Kristen Delevich Assistant Professor, Washington State University
Cinthya Zepeda Mendoza Medical Director, Cytogenetics and Genomic Microarray at ARUP Laboratories. Assistant Professor, Department of Pathology, University of Utah
Wee Siong Goh Assistant Professor Shenzhen Bay Laboratory, China
Arkarup Banerjee Assistant Professor, Cold Spring Harbor Laboratory, NY
Charles Underwood Group Leader, Max Planck Institute for Plant Breeding Research, Cologne
Justus Kebschull Assistant Professor, The Johns Hopkins University

Graduates in Industry Positions (in order of completion)

Student Current Position
Amy Caudy Principal, Scientific Operations, Maple Flavored Solutions, NY*
Emiliano Rial-Verde Vice President, Food & Ingredients Strategy, Bunge Limited, NY
Rebecca Ewald International Business Leader, Ventana Medical Systems/Roche, AZ
Catherine Seiler (nee Cormier) Head, Biomarker Operations, H3 Medicine, MA
Yaniv Erlich Chief Science Officer, My Heritage, Israel*
Colin Malone Co-Founder & Head of Biology at VNV NewCo, NY*
Amy Rappaport Senior Scientist, Gristone Oncology, CA
Frederick Rollins Director, Competitive Intelligence, Oncology R&D, AstraZeneca, Washington DC
Patrick Finigan Associate Director, Regulatory Affairs CMC, Gilead Sciences
Kyle Honegger Data Scientist, Ann & Robert H. Lurie Children’s Hospital of Chicago, IL
Maria Pineda Co-Founder, CEO, Envisagenics, NY
Felix Schlesinger Bioinformatics Scientist, Illumina, Inc. San Diego, CA
Megan Hogan (nee Bodnar) Senior Scientist, Neochromosome, Inc., NY
Paloma Guzzardo Head, Target Biology, Variant Bio, WA
Saya Ebbesen Associate Director, Medical + Scientific Strategy at BluPrint Oncology, London, UK
Marek Kudla Software developer, Design Everest
Joshua Sanders Founder and C.E.O., Sanworks, L.L.C., NY
Katie Liberatore Senor Manager, Trait Discovery & Targeting, Calyxt, Minneapolis, MN
Kaja Wasik Co-Founder and Chief Scientific Officer, Gencove & Variant Bio, NY
Stephane Castel Co-Founder at Variant Bio and Senior Research Fellow at New York Genome Center
Mitchell Bekritsky Senior Bioinformatics Scientist, Illumina, Inc. Cambridge, UK
Sang-Geol Koh Scientist and Entrepreneur, {Mind}, South Korea
Ozlem Aksoy (nee Mert) Senior Scientist, Pfizer, CA
Susann Weissmueller Strategic Partnering Associate, Roche, Switzerland
Nilgun Tasdemir Senior Scientist, Pfizer, NY
Silvia Fenoglio Senior Scientist, Tango Therapeutics, MA
Jack Walleshauser Senior Scientist Revolution Medicines, CA
Lisa Krug Scientist, Kallyope, NY
Robert Aboukhalil Senior Software Engineer, Chan Zuckerberg Initiative
Anja Hohmann Associate Director, Discovery Biology, Constellation Pharmaceuticals, MA
Matt Koh Research Engineer, ASAAP, NY
Annabel Romero Hernandez Scientist, Regeneron Pharmaceuticals, NY
Maria Nattestad Senior Software Engineer, Google, CA
Onyekachi Odoemene Principal Data Scientist, Capital One, MD
Daniel Kepple Senior Machine Learning Engineer, Samsung Artificial Intelligence Center, NY
Kristina Grigaityte Senior Data Scientist, Tempus Labs, Inc, NY
Elizabeth Hutton Computational Biologist, ArtisanBio, CO
Ngoc (Tumi) Tran Machine Learning Scientist, Relay Therapeutics, MA
Shaina Lu Bioinformatics Scientist, Eagle Genomics
Benjamin Harris Computational Biologist, Lyell Immunopharma, CA

*Left a faculty position

Graduates in Postdoctoral or Academic Research Positions (in order of completion)

Student Current Position
Charles Kopec Associate Professional Specialist, Princeton University (Advisor, Dr. Carlos Brody)
Oliver Tam Computational Science Analyst, Cold Spring Harbor Laboratory, NY
Galen Collins Postdoctoral Fellow, Harvard Medical School (Advisor: Dr. Alfred Goldberg)
David Simpson Postdoctoral Fellow, Stanford University (Advisor: Dr. Alejandro Sweet-Cordero)
Claudio Scuoppo Instructor, Columbia University (Advisor: Riccardo Dalla-Favera)
Zhenxun Wang Research Fellow, Genome Institute of Singapore with (Advisor: Wai Leong Tam)
Eyal Gruntman Research Scientist, Janelia Farm (Advisor: Michael Reiser)
Yevgeniy Plavskin Postdoctoral Fellow, New York University (Advisor: Mark Siegal)
Hassana Oyibo Viral Vector Core Lead Scientist, Friedrich Miescher Institute for Biomedical Research, Switzerland
Zinaida Perova Project Lead, Cancer Models, European Bioinformatics Institute (EMBL-EBI), Cambridge, UK
Dario Bressan Head of IMAXT laboratory, CRUK, Cambridge Institute, UK
Philippe Batut Postdoctoral Fellow, Princeton University (Advisor: Michael Levine)
Elvin Wagenblast Postdoctoral Fellow, University of Toronto (Advisor: John Dick)
Joaquina Delas Vives Postdoctoral Fellow, Francis Crick Institute, UK (Advisor: James Briscoe)
Abram Santana Postdoctoral Fellow, Harvard University (Advisor: Joan Brugge)
Fred Marbach Postdoctoral Fellow, Sainsbury Wellcome Centre, UK (Advisor: Marcus Stephenson-Jones)
Yu-Jui (Ray) Ho Computational Biologist, Memorial Sloan-Kettering Cancer Center (Advisor: Scott Lowe)
Paul Masset Postdoctoral Fellow, Harvard University (Advisor: Venkatesh Murthy and Naoshige Uchida)
Talitha Forcier Postdoctoral Fellow, Cold Spring Harbor Laboratory (Advisor: Molly Hammell)
Laura Maiorino Postdoctoral Fellow, Koch Institute, MIT (Advisor: Darrell Irvine)
Michael Gutbrod Postdoctoral Fellow, MIT (Advisor: Manolis Kellis)
Giorgia Battistoni Research Associate, Cancer Research UK (Advisor: Gregory Hannon)
Sashank Pisupati Postdoctoral Fellow, Princeton University (Advisor: Yael Niv)
Jacqueline Giovanniello Postdoctoral Fellow, UCLA (Advisor: Kate Wassum)
Hamza Giaffar Postdoctoral Fellow, UCSD (Advisor: Mikio Aoi)
Anqi Zhang Postdoctoral Fellow, Harvard University (Advisor: Florian Engert)
Alexandra Nowlan Postdoctoral Fellow, UNC Chapel Hill (Advisor: Zoe McElligott)
Katie Meze Postdoctoral Fellow, University of Michigan (Advisor: Nils Walters)
Brianna Bibel Postdoctoral Fellow, UCSF (Advisor: Danica Fujimori)

Graduates in non-Research Positions (in order of completion)

Name Current Position
Ahmet M. Denli Associate Editor, Genome Research, CSHL, NY
Beth Chen Operations Manager at Homer Scientific Holdings Inc.
Darren Burgess Senior Editor, Nature Reviews Genetics, UK
Rebecca Bish-Cornelissen Scientific Director, The Mark Foundation for Cancer Research, NY
Angelique Girard Director of Finance and Administration, Amplitude Studios, Paris, France
Allison Blum Managing Director, Scientific Communications, Life Sci Advisors, NY
Keisha John Associate Dean for Diversity, Equity and Inclusion, University of Virginia, VA
Elizabeth Nakasone Heme-Onc Fellow, Fred Hutchinson Cancer Research Center
Ian Peikon Venture Partner, Lux Capital, NY
Colleen Carlston Clinical Fellow in Pediatrics, Boston Children’s Hospital, MA
Tyler Garvin VP Operations, Underground Cellar, CA
Brittany Cazakoff Associate Lawyer, Cooley, CA
Lital Chartarifsky Associate Medical Director, FCB CURE, NY
Michael Gutbrod Associate, SANTÉ
Emilis Bruzas Medical Director, FCB Health, NY
Sanchari Ghosh Content Specialist for BioRxiv, NY
Georgi Yordanov Equity Research Associate, Cowen and Company, NY
Jue Xiang Wang Consultant, Boston Consulting Group, NY
Matt Lee Clinical Science Liaison, TG Therapeutics
Sofya Polyanskaya Consultant, Scitaris, Berlin, Germany