Since the School’s first graduating class in 2004, 145 students have been awarded doctoral degrees from the School. The overall retention rate is 95%. Of those earning a degree: 89% earned a Ph.D.; 11% graduated with a Master’s degree.
All students complete their degree within 6 years. The average time to degree is 5.19 years.
Students have published almost 500 papers, a demonstration of the rigorous and reproducible work they performed during their training. About a quarter of the students have successfully applied for external fellowships.
A third of our graduates are in faculty positions, most in tenure-track positions at major U.S or foreign research institutions, and a third in industry. Other graduates are in academic research, postdoctoral research, or other scientific careers, such as university administrators, CEOs of biotech companies, and consultants.
Ph.D. Student Progress & Time to Degree
Career Paths of Ph.D. Students
Graduates in Faculty or Independent Positions (in order of completion)
| Name | Faculty Position |
|---|---|
| Ira Hall | Professor, Yale University, CT |
| Niraj Tolia | Senior Investigator, NIAID, National Institutes of Health |
| Patrick Paddison | Professor, Fred Hutchinson Cancer Research Center, WA |
| Elizabeth Bartom (nee Thomas) | Assistant Professor, Northwestern University, IL |
| Michelle Heck (nee Cilia) | Research Molecular Biologist, USDA-ARS and Adjunct Associate Professor, Cornell University |
| Zachary Lippman | Professor, Cold Spring Harbor Laboratory & Investigator, Howard Hughes Medical Institute |
| Ji-Joon Song | Professor, Korea Advanced Institute of Science and Technology (KAIST), South Korea |
| Elena Ezhkova | Associate Professor, Mount Sinai School of Medicine, NY |
| Masafumi Muratani | Professor, University of Tsukuba, Japan |
| Santanu Chakraborty | Program Director for Sustainable Life Sciences and Associate Professor, Atria University, India |
| Claudia Feierstein | Research Associate, Champalimaud Neuroscience Programme, Portugal |
| Gowan Tervo | Group Leader, HHMI, Janelia Research Campus, VA |
| Marco Mangone | Professor, Arizona State University, AZ |
| Elizabeth Murchison | Professor, Cambridge University, UK |
| Hiroki Asari | Group leader, EMBL Monterotondo, Rome |
| François Bolduc | Associate Professor, University of Alberta, Canada |
| Wei Wei | Professor, University of Chicago, IL |
| Christopher Harvey | Associate Professor, Harvard University, MA |
| Tomas Hromadka | Group Leader, Institute of Neuroimmunology, Slovak Academy of Sciences |
| Monica Dus | Associate Professor, University of Michigan, MI and White House Fellow, United States Department of the Navy |
| Shu-Ling Chiu | Assistant Professor, Academia Sinica, Taiwan |
| Daniel Chitwood | Assistant Professor, Michigan State University, MI |
| Jeremy Wilsuz | Associate Professor, Baylor College of Medicine, TX |
| Shraddha Pai | Investigator, Ontario Institute for Cancer Research, Toronto |
| Oliver Fregoso | Assistant Professor, University of California, Los Angeles, CA |
| Hiroshi Makino | Assistant Professor, Nanyang Technological University, Singapore |
| Galen Collins | Assistant Professor, Mississippi State University, MS |
| Katherine McJunkin | Stadtman Tenure Track Investigator, National Institutes of Health |
| Zhenxun Wang | Adjunct Junior Principal Investigator, Singapore Eye Research Institute and Adjunct Assistant Professor Duke-NUS Medical School, Singapore |
| Eyal Gruntman | Assistant Professor, University of Toronto Scarborough, Canada |
| Petr Znamenskiy | Assistant Professor, The Crick Institute, UK |
| Yevgeniy Plavskin | Visiting Clinical Associate Professor, New York University |
| Michael Pautler | Head, Genomics Services, Platform Genetics & Vineland Research and Innovation Centre, Canada |
| Melanie Eckersley-Maslin | Assistant Professor, Peter Mac Cancer Centre at University of Melbourne |
| Philippe Batut | Assistant Professor, Columbia University (January 2024) |
| Elvin Wagenblast | Assistant Professor, Mount Sinai School of Medicine, NY |
| Kristen Delevich | Assistant Professor, Washington State University |
| Cinthya Zepeda Mendoza | Medical Director, Cytogenetics and Genomic Microarray at ARUP Laboratories. Assistant Professor, Department of Pathology, University of Utah |
| Wee Siong Goh | Assistant Professor Shenzhen Bay Laboratory, China |
| Arkarup Bandyopadhyay | Assistant Professor, Cold Spring Harbor Laboratory, NY |
| Charles Underwood | Professor, Radboud University, The Netherlands |
| Justus Kebschull | Assistant Professor, The Johns Hopkins University |
| Paul Masset | Assistant Professor, McGill University, Canada (January 2024) |
| Brianna Bibel | Visiting Professor, Saint Mary’s College of California |
Graduates in Industry Positions (in order of completion)
| Student | Current Position |
|---|---|
| Amy Caudy | Principal, Scientific Operations, Maple Flavored Solutions, NY* |
| Emiliano Rial-Verde | Vice President, Food & Ingredients Strategy, Bunge Limited, NY |
| Rebecca Ewald | International Business Leader, Ventana Medical Systems/Roche, AZ |
| Catherine Seiler (nee Cormier) | Director, Operational Excellence in Translation Medicine, AstraZeneca, MA |
| Allison Blum | Senior Director, Scientific Affairs, Pfizer Research and Development, NY |
| Yaniv Erlich | CEO, Eleven Therapeutics, UK* |
| Colin Malone | Co-Founder & Head of Biology at VNV NewCo, NY* |
| Amy Leung | Principal Solution Engineer, DNAnexus, CA |
| Amy Rappaport | Senior Director, Translational Immunology, Gritstone bio, Inc., CA |
| Claudio Scuoppo | Principal Scientist, Sapience Therapeutics, Inc. |
| Frederick Rollins | Director, Competitive Intelligence, Oncology R&D, AstraZeneca, Washington DC |
| Patrick Finigan | Associate Director, Regulatory Affairs CMC, Gilead Sciences |
| Kyle Honegger | Data Scientist, Ann & Robert H. Lurie Children’s Hospital of Chicago, IL |
| Maria Pineda | Co-Founder, CEO, Envisagenics, NY |
| Felix Schlesinger | Bioinformatics Lead, Scale Bio, CA |
| Megan Hogan (nee Bodnar) | Team Lead, Neochromosome, Inc., NY |
| Paloma Guzzardo | Head, Target Biology, Variant Bio, WA |
| Saya Ebbesen | Associate Director, Medical + Scientific Strategy at BluPrint Oncology, London, UK |
| Hassana Oyibo | Associate Director, Synthetic Biology, Tessera Therapeutics |
| Michael Pautler | Head, Genomics Services, Platform Genetics, Canada* |
| Marek Kudla | Director of Bioinformatics, Ardigen, Poland |
| Joshua Sanders | Founder and C.E.O., Sanworks, L.L.C., NY |
| Katie Liberatore | Molecule Design Director, Cibus, MN |
| Kaja Wasik | Co-Founder and Chief Scientific Officer, Gencove & Variant Bio, NY |
| Stephane Castel | Co-Founder and Chief Technology Officer, Variant Bio, WA |
| Mitchell Bekritsky | Senior Manager, Bioinformatics, Illumina, Inc., |
| Sang-Geol Koh | Scientist and Entrepreneur, {Mind}, South Korea |
| Ozlem Aksoy (nee Mert) | Senior Scientist, Pfizer, CA |
| Susann Manchado (nee Weissmueller) | Strategic Assistant (Chief of Staff) for President Global Drug Development, Novartis, Switzerland |
| Nilgun Tasdemir | Senior Scientist, Pfizer, NY |
| Silvia Fenoglio | Senior Scientist, Tango Therapeutics, MA |
| Jack Walleshauser | Senior Scientist, Spotlight Therapeutics, CA |
| Lisa Krug | Senior Scientist, Kallyope, NY |
| Robert Aboukhalil | Senior Software Engineer, Chan Zuckerberg Initiative |
| Anja Hohmann | Director, Cell Therapies at Be Biopharma, MA |
| Matt Koh | Research Engineer, ASAAP, NY |
| Annabel Romero Hernandez | Director of AI for Drug Discovery and Molecular Modeling, SFL Scientific, MA |
| Maria Nattestad | Senior Software Engineer, Google, CA |
| Onyekachi Odoemene | Research Scientist, ML Solutions Lab, Amazon |
| Daniel Kepple | Engineer, Meta, NY |
| Sashank Pisupati | Machine Learning Scientist, Limbic, UK |
| Kristina Grigaityte | Associate Director, Data Science, Merck |
| Elizabeth Hutton | Senior Computational Biologist, ArtisanBio, CO |
| Ngoc (Tumi) Tran | Machine Learning Scientist, Relay Therapeutics, MA |
| Katie Meze | Senior Scientist, Structural Biology, Sanofi, MA |
| Benjamin Berube | Research Scientist, Monocot Breeding and Biotechnology, Ohalo Genetics, CA |
| Shaina Lu | Senior Data Scientist, R&D, Miltenyi Biotec |
| Benjamin Harris | Computational Biologist II, Lyell Immunopharma, CA |
| Christopher Krasniak | Research Data Scientist, Somatus, NH |
| Ran (Rena) Yan | Senior Scientist, Parthenon Therapeutics, MA |
| Martyna Sroka | Scientist, Relay Therapeutics, MA |
| Sara Boyle | Data Analytics Consultant, Northern Trust, IL |
| Asad Aziz Lakhani | Associate, Search and Evaluation, Autobahn Labs |
*Left a faculty position
Graduates in Postdoctoral or Academic Research Positions (in order of completion)
| Student | Current Position |
|---|---|
| Charles Kopec | Associate Professional Specialist, Princeton University (Advisor, Dr. Carlos Brody) |
| Oliver Tam | Research Scientist, NYU Institute for Systems Genetics, NY |
| Elizabeth Nakasone | Junior Researcher, Translational and Clinical Research, University of Hawaiʻi Cancer Center |
| Claudio Scuoppo | Instructor, Columbia University (Advisor: Riccardo Dalla-Favera) |
| Zhenxun Wang | Research Fellow, Genome Institute of Singapore with (Advisor: Wai Leong Tam) |
| Eyal Gruntman | Research Scientist, Janelia Farm (Advisor: Michael Reiser) |
| Yevgeniy Plavskin | Postdoctoral Fellow, New York University (Advisor: Mark Siegal) |
| Zinaida Perova | Project Lead, Cancer Models, European Bioinformatics Institute (EMBL-EBI), Cambridge, UK |
| Dario Bressan | Head of IMAXT laboratory, CRUK, Cambridge Institute, UK |
| Philippe Batut | Postdoctoral Fellow, Princeton University (Advisor: Michael Levine) |
| Joaquina Delas Vives | Postdoctoral Fellow, Francis Crick Institute, UK (Advisor: James Briscoe) |
| Abram Santana | Postdoctoral Fellow, Harvard University (Advisor: Joan Brugge) |
| Fred Marbach | Senior Laboratory Research Scientist, Francis Crick Institute, UK (Advisor: Andreas Schaefer) |
| Yu-Jui (Ray) Ho | Bioinformatic Specialist II, Memorial Sloan-Kettering Cancer Center (Advisor: Scott Lowe) |
| Paul Masset | Postdoctoral Fellow, Harvard University (Advisor: Venkatesh Murthy and Naoshige Uchida) |
| Talitha Forcier | Postdoctoral Fellow, New York University (Advisor: Molly Hammell) |
| Laura Maiorino | Postdoctoral Fellow, Koch Institute, MIT (Advisor: Darrell Irvine) |
| Giorgia Battistoni | Research Associate, Cancer Research UK (Advisor: Gregory Hannon) |
| Jacqueline Giovanniello | Postdoctoral Fellow, UCLA (Advisor: Kate Wassum) |
| Hamza Giaffar | Postdoctoral Fellow, UCSD (Advisor: Mikio Aoi) |
| Anqi Zhang | Postdoctoral Fellow, Harvard University (Advisor: Florian Engert) |
| Alexandra Nowlan | Postdoctoral Fellow, UNC Chapel Hill (Advisor: Zoe McElligott) |
| Katie Meze | Postdoctoral Fellow, University of Michigan (Advisor: Nils Walters) |
| Christopher Krasniak | Data Science Fellow, The Data Incubator |
| Luqun Shen | Postdoctoral Fellow, Cornell University (Advisor: Alex Kwan) |
| Alberto Corona | Postdoctoral Fellow, Icahn School of Med at Mount Sinai (Advisor: Paul Kenny) |
| Kathryn O’Neill | Child Health Equity Postdoc, University of Washington, Seattle |
| David Johnson | Postdoctoral Fellow, Invent at Seattle Children’s Postdoctoral Scholars Program (Advisor: Siobhan Pattwell) |
| Bruno Gegenhuber | Postdoctoral Fellow, Harvard University (Advisor: Michael Greenberg) |
| Lyndsey Aguirre | Biological Sciences Tech, Peanuts and Small Grains Research Unity, USDA-ARS, OK |
| Amritha Varshini Hanasoge Somasundara | Senior Scientist, Memorial Sloan-Kettering Cancer Center (Advisor: Ross Levine) |
| Chengxiang (Charlie) Yuan | Postdoctoral Fellow, Institute of Molecular and Cell Biology, Singapore (Advisor: Weimiao Yu) |
| Diogo Maia e Silva | Postdoctoral Fellow, Mass General Hospital (Advisor: Keith Joung) |
| Yuzhao (Richard) Hu | Postdoctoral Scientist, Corteva Agriscience, IA |
Graduates in non-Research Positions (in order of completion)
| Name | Current Position |
|---|---|
| Ahmet M. Denli | Associate Editor, Genome Research, CSHL, NY |
| Beth Chen | Operations Manager at Homer Scientific Holdings Inc. |
| Darren Burgess | Senior Editor, Nature Reviews Genetics, UK |
| Rebecca Bish-Cornelissen | Head of Discovery and Preclinical Research, The Mark Foundation for Cancer Research, NY |
| Angelique Girard | Director of Finance and Administration, Amplitude Studios, Paris, France |
| Allison Blum | Managing Director, Scientific Communications, Life Sci Advisors, NY |
| Keisha John | Associate Dean for Diversity, Equity and Inclusion, University of Virginia, VA |
| David Simpson | Freelance Data Scientist and Strategy Consultant |
| Ian Peikon | Co-founder & CSO, Cajal Neuroscience; Venture Partner, Lux Capital |
| Colleen Carlston | Clinical Fellow in Pediatrics, Boston Children’s Hospital, MA |
| Tyler Garvin | VP Operations, Underground Cellar, CA |
| Brittany Cazakoff | Associate Lawyer, Cooley, CA |
| Lital Chartarifsky | Associate Medical Director, FCB CURE, NY |
| Michael Gutbrod | Senior Associate, Santé Ventures, MA |
| Emilis Bruzas | Associate Director, Medical Strategy, FCB Health, NY |
| Sanchari Ghosh | Content Specialist for BioRxiv, NY |
| Georgi Yordanov | Equity Research Associate, Cowen and Company, NY |
| Jue Xiang Wang | Consultant, Boston Consulting Group, NY |
| Matt Lee | Oncology Medical Science Liaison, EMD Serono, Inc. |
| Sofya Polyanskaya | Consultant, Scitaris, Berlin, Germany |
| Jenelys Ruiz Ortiz | Bioinformatics Analyst, Bioinformatics Resource Center, The Rockefeller University, NY |
| Ilgin Ergin | Foundation and Corporate Relations Coordinator, CSHL, NY |
| Alexa Pagliaro | Foundation and Corporate Relations Officer, Whitehead Institute, MA |